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0 History Of Shakespeare

William Shakespeare was baptised in Stratford-Upon-Avon, Warwickshire, on April 26, 1564. April 23, 1564 is the day that most believe to be his birthday, but there are no records that state what day he was born. His father, John Shakespeare, was a whittawer by profession and held several important town offices. His mother, Mary Arden, was from a fairly wealthy family. In all the Shakespeares had eight children, and William was their first son.Shakespeare did not have an extensive education. He attended Stratford grammar school, a school that specifically served prominent citizens. It is not known how long Shakespeare attended, but it is assumed that this is the only education he recieved.
On November 28, 1582 Shakespeare married Anne Hathaway, a farmer's daughter. Anne was twenty six years old when they married, and William was only eighteen. Anne and William's first daughter, Susanna, was baptised on May 26, 1582. A set of twins, Hamnet and Judith, were born two years later. In 1596 Hamnet died at the age of eleven. Between 1585 and 1592 no records of Shakespeare were found, and that period of his life is usually referred to as "The Lost Years". Some have speculated that he either became a school teacher, became a butcher's apprentice, or was running from the law during this time.
The first evidence of Shakespeare after 1592 was in London. Here he had established himself as a playwright and actor and had found a sponsor, Henry Wriothsley. However, Shakespeare's work in the theaters came to a halt in January of 1593 when the theaters closed because of the plague. The company that Shakespeare worked for was called "Lord Chamberlain's Men" and changed their name to "The King's Men" after King James I took over in 1603. Because Shakespeare worked and performed for them, this company became the biggest and most famous acting company. Shakespeare became very wealthy as a director, writer, actor, and stockholder in "The King's Men".
In 1611 Shakespeare retired and left London. He made a will on March 25, 1616, and died on April 23, 1616. He was fifty two years old. The cause of Shakespeare's death is not known. Shakespeare also wrote his own epitaph because during his time, when the graveyard was full, people would dig up someone's corpse and burn it so that another could be buried in that person's place. This disgusted Shakespeare, and he didn't want this type of disrespect after his death. His epitaph reads as follows:
"Good Friends, for Jesus' sake forbear,
To dig the bones enclosed here!
Blest be the man that spares these stones,
And curst be he that moves my bones."
To this day no one has disturbed Shakespeare's grave.
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0 History Of Halloween

Historian Nicholas Rogers, exploring the origins of Halloween, notes that while "some folklorists have detected its origins in the Roman feast of Pomona, the goddess of fruits and seeds, or in the festival of the dead called Parentalia, it is more typically linked to the Celtic festival ofSamhain, whose original spelling was Samuin (pronounced sow-an or sow-in)". The name of the festival historically kept by the Gaels in the British isles is derived from Old Irish and means roughly "summer's end".
The word Halloween is first attested in the 16th century and represents a Scottish variant of the fuller All-Hallows-Even ("evening"), that is, the night before All Hallows Day. Although the phrase All Hallows is found in Old English (ealra hālgena mæssedæg, mass-day of all saints), All-Hallows-Even is itself not attested until 1556.

Development of artifacts and symbols associated with Halloween formed over time. For instance, the carving of jack-o'-lanterns springs from the souling custom of carving turnips into lanterns as a way of remembering the souls held in purgatory. The turnip has traditionally been used in Ireland and Scotland at Halloween, but immigrants to North America used the native pumpkin, which are both readily available and much larger – making them easier to carve than turnips. The American tradition of carving pumpkins is recorded in 1837 and was originally associated with harvest time in general, not becoming specifically associated with Halloween until the mid-to-late 19th century.
The imagery of Halloween is derived from many sources, including national customs, works of Gothic andhorror literature (such as the novels Frankenstein and Dracula), and classic horror films (such asFrankenstein and The Mummy). Among the earliest works on the subject of Halloween is from Scottish poet John Mayne in 1780, who made note of pranks at Halloween; "What fearfu' pranks ensue!", as well as the supernatural associated with the night, "Bogies" (ghosts), influencing Robert BurnsHalloween 1785.Elements of the autumn season, such as pumpkins, corn husks, and scarecrows, are also prevalent. Homes are often decorated with these types of symbols around Halloween.
Halloween imagery includes themes of deathevil, the occult, or mythical monsters. Black and orange are the holiday's traditional colors.

Trick-or-treating is a customary celebration for children on Halloween. Children go in costume from house to house, asking for treats such as candy or sometimes money, with the question, "Trick or treat?" The word "trick" refers to a (mostly idle) "threat" to perform mischief on the homeowners or their property if no treat is given. In some parts of Scotland children still go guising. In this custom the child performs some sort of trick, i.e. sings a song or tells a ghost story, to earn their treats.
The practice of dressing up in costumes and begging door to door for treats on holidays dates back to theMiddle Ages and includes Christmas wassailing. Trick-or-treating resembles the late medieval practice ofsouling, when poor folk would go door to door on Hallowmas (November 1), receiving food in return for prayers for the dead on All Souls' Day (November 2). It originated in Ireland and Britain, although similar practices for the souls of the dead were found as far south as Italy. Shakespeare mentions the practice in his comedy The Two Gentlemen of Verona (1593), when Speed accuses his master of "puling [whimpering or whining] like a beggar at Hallowmas."
In Scotland and Ireland, Guising — children disguised in costume going from door to door for food or coins — is a traditional Halloween custom, and is recorded in Scotland at Halloween in 1895 where masqueraders in disguise carrying lanterns made out of scooped out turnips, visit homes to be rewarded with cakes, fruit and money. The practise of Guising at Halloween in North America is first recorded in 1911, where a newspaper inKingston, Ontario reported children going "guising" around the neighborhood.
The taste in Hallowe'en festivities now is to study old traditions, and hold a Scotch party, using Burn's poem Hallowe'en as a guide; or to go a-souling as the English used. In short, no custom that was once honored at Hallowe'en is out of fashion now.
American historian and author Ruth Edna Kelley of Massachusetts wrote the first book length history of the holiday in the U.S; The Book of Hallowe'en (1919), and references souling in the chapter "Hallowe'en in America";
In her book, Kelley touches on customs that arrived from across the Atlantic; "Americans have fostered them, and are making this an occasion something like what it must have been in its best days overseas. All Hallowe'en customs in the United States are borrowed directly or adapted from those of other countries".
While the first reference to "guising" in North America occurs in 1911, another reference to ritual begging on Halloween appears, place unknown, in 1915, with a third reference in Chicago in 1920.
Hallowe’en provided an opportunity for real strenuous fun. No real damage was done except to the temper of some who had to hunt for wagon wheels, gates, wagons, barrels, etc., much of which decorated the front street. The youthful tormentors were at back door and front demanding edible plunder by the word “trick or treat” to which the inmates gladly responded and sent the robbers away rejoicing.
The earliest known use in print of the term "trick or treat" appears in 1927, from Blackie, Alberta, Canada:
The thousands of Halloween postcards produced between the turn of the 20th century and the 1920s commonly show children but do not depict trick-or-treating. The editor of a collection of over 3,000 vintage Halloween postcards writes, "There are cards which mention the custom [of trick-or-treating] or show children in costumes at the doors, but as far as we can tell they were printed later than the 1920s and more than likely even the 1930s. Tricksters of various sorts are shown on the early postcards, but not the means of appeasing them".Trick-or-treating does not seem to have become a widespread practice until the 1930s, with the first U.S. appearances of the term in 1934,and the first use in a national publication occurring in 1939.

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0 History Of The Beatles


The Beatles were an English rock band, formed in Liverpool in 1960. They are one of the most commercially successful and critically acclaimed acts in the history of popular music. From 1962, the group consisted of John Lennon (rhythm guitar, vocals), Paul McCartney (bass guitar, vocals), George Harrison (lead guitar, vocals) and Ringo Starr (drums, vocals). Rooted in skiffleand 1950s rock and roll, the group later worked in many genres ranging from pop ballads topsychedelic rock, often incorporating classical and other elements in innovative ways. The nature of their enormous popularity, which first emerged as "Beatlemania", transformed as their songwriting grew in sophistication. They came to be perceived as the embodiment of ideals of the social and cultural revolutions of the 1960s.
Initially a five-piece line-up of Lennon, McCartney, Harrison, Stuart Sutcliffe (bass) and Pete Best(drums), they built their reputation playing clubs in Liverpool and Hamburg over a three-year period from 1960. Sutcliffe left the group in 1961, and Best was replaced by Starr the following year. Moulded into a professional outfit by their manager, Brian Epstein, their musical potential was enhanced by the creativity of producer George Martin. They achieved mainstream success in the United Kingdom in late 1962, with their first single, "Love Me Do". Gaining international popularity and acquiring the nickname "Fab Four" the following year, they toured extensively until 1966. The band then retreated to recording studios until their break-up in 1970. Each subsequently found success in independent musical careers.
During their studio years, they produced what critics consider some of their finest material including the album Sgt. Pepper's Lonely Hearts Club Band (1967), widely regarded as a masterpiece. They are the best-selling band in the history of popular music, and over four decades after their break-up, their recordings are still in demand. They have had more number one albums on the UK charts and have held the top spot longer than any other musical act. According to the RIAA, they have sold more albums in the United States than any other artist. The Beatles were placed at number one on Billboard magazine's fiftieth-anniversary list of all-time top Hot 100 artists in 2008. They have received 7 Grammy Awards from the AmericanNational Academy of Recording Arts and Sciences and 15 Ivor Novello Awards from the British Academy of Songwriters, Composers and Authors. They were collectively included in Timemagazine's compilation of the 20th century's 100 most influential people. Lennon wasmurdered outside his home in New York City in 1980, and Harrison died of cancer in 2001. McCartney and Starr remain active


Formation and early years (1957–1962)

Aged sixteen, John Lennon formed a skiffle group called The Quarrymen, in March 1957. Fifteen-year-old Paul McCartney joined as a guitarist after he and Lennon met that July. When McCartney in turn invited George Harrison to watch the group the following February, the fourteen-year-old joined as lead guitarist. By 1960, Lennon's schoolfriends had left the group, he had begun studies at the Liverpool College of Art and the three guitarists were playing rock and rollwhenever they could get a drummer. Lennon's fellow student Stu Sutcliffe joined on bass in January. Sutcliffe suggested changing the band name to "The Beetles" as a tribute to Buddy Hollyand The Crickets, and they became "The Beatals" for the first few months of the year. In May 1960 they undertook a brief tour of Scotland, as backing group for pop singer Johnny Gentle. After trying other names including "Johnny and the Moondogs", "Long John and The Beetles" and "The Silver Beatles", the band finally became "The Beatles" in August. The lack of a permanent drummer posed a problem when the group's unofficial manager, Allan Williams, arranged a resident band booking for them in Hamburg, Germany. Before the end of August they auditioned and hiredPete Best, and the five-piece band left for Hamburg four days later, contracted to club ownerBruno Koschmider, for a 48-night residency. "Hamburg in those days did not have rock 'n' roll music clubs. It had strip clubs", says Beatles biographer Philip Norman.
Bruno had the idea of bringing in rock groups to play in various clubs. They had this formula. It was a huge nonstop show, hour after hour, with a lot of people lurching in and the other lot lurching out. And the bands would play all the time to catch the passing traffic. In an American red-light district, they would call it nonstop striptease.
Many of the bands that played in Hamburg were from Liverpool. ... It was an accident. Bruno went to London to look for bands. But he happened to meet a Liverpool entrepreneur in Soho, who was down in London by pure chance. And he arranged to send some bands over.
Harrison, only 17 years old in August 1960, obtained permission to stay in Hamburg by lying to the German authorities about his age. Initially placing the group at the Indra Club, Koschmider moved them to the Kaiserkeller in October after the Indra was closed down due to noise complaints. When they violated their contract by performing at The Top Ten Club, a rival venue, Koschmider reported the underage Harrison to the authorities, leading to his deportation in November. A week later, McCartney and Best were arrested for arson after they set fire to a condom nailed to a wall in their room; they were also deported. Lennon returned to Liverpool in mid-December, while Sutcliffe remained in Hamburg with his new German fiancée, Astrid Kirchherr, for another month. Kirchherr took the first professional photos of the group and cut Sutcliffe's hair in the German "exi" (existentialist) style of the time, a look later adopted by the other Beatles.
In Liverpool, the Merseybeat movement was gathering force. The band had their first recording session under Martin's direction at EMI's Abbey Road Studios in London in June 1962. Martin complained to Epstein about Best's drumming and suggested the band use a session drummer in the studio. Instead, Best was replaced by Ringo Starr. Starr, who left Rory Storm and the Hurricanes to join The Beatles, had already performed with them during Best's occasional absences. Martin still hired session drummer Andy White for one session. White played on the single "Love Me Do" and "P.S. I Love You". Released in October, "Love Me Do" was a top twenty UK hit, peaking at number seventeen on the chart. After a November studio session that yielded what would be their second single, "Please Please Me", they made their TV debut with a live performance on the regional news programme People and Places.
During the next two years, the group were resident for further periods in Hamburg. They used Preludin both recreationally and to maintain their energy through all-night performances. When Sutcliffe decided to leave the band in early 1961 and resume his art studies in Germany, McCartney took up the bass. German producer Bert Kaempfert contracted what was now a four-piece to act as Tony Sheridan's backing band on a series of recordings. Credited to "Tony Sheridan & The Beat Brothers", the single "My Bonnie", recorded in June and released four months later, reached number 32 on the Musikmarkt chart. The Beatles were also becoming more popular back home in Liverpool. During one of the band's frequent appearances there at the Cavern Club, they encountered Brian Epstein, a local record store owner and music columnist. When the band appointed Epstein manager in January 1962, Kaempfert agreed to release them from the German record contract. After Decca Records rejected the band with the comment "Guitar groups are on the way out, Mr. Epstein",George Martin signed the group to EMI's Parlophone label. News of a tragedy greeted them on their return to Hamburg in April. Meeting them at the airport, a stricken Kirchherr told them of Sutcliffe's death from a brain haemorrhage.
The band concluded their last Hamburg stint in December 1962. By now it had become the pattern that all four members contributed vocals, although Starr's restricted range meant he rarely sang lead. Lennon and McCartney had established a songwriting partnership; as the band's success grew, their celebrated collaboration limited Harrison's opportunities as a lead vocalist. Epstein, sensing their commercial potential, encouraged the group to adopt a professional attitude to performing. Lennon recalled the manager saying, "Look, if you really want to get in these bigger places, you're going to have to change—stop eating on stage, stop swearing, stop smoking." Lennon said, "We used to dress how we liked, on and off stage. He'd tell us that jeans were not particularly smart and could we possibly manage to wear proper trousers, but he didn't want us suddenly looking square. He'd let us have our own sense of individuality ... it was a choice of making it or still eating chicken on stage."
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0 History Of Dinasaurs


Dinosaurs (from Greekδεινός terrible or potent, and σαύρα lizard) are a diverse group of animals that were the dominant terrestrial vertebrates for over 160 million years, from the late Triassicperiod (about 230 million years ago) until the end of the Cretaceous (about 65 million years ago), when the Cretaceous–Paleogene extinction event led to the extinction of most dinosaur species at the close of the Mesozoic era. The fossil record indicates that birds evolved within theropoddinosaurs during the Jurassic period. Some of them survived the Cretaceous–Paleogene extinction event, including the ancestors of all modern birds. Consequently, in modern classification systems, birds are considered a type of dinosaur—the only group of which that has survived to the present day.
Dinosaurs are a diverse and varied group of animals; birds, at over 9,000 species, are the most diverse group of vertebrate besides perciform fish. Paleontologists have identified over 500 distinct genera and more than 1,000 different species of non-avian dinosaurs. Dinosaurs are represented on every continent by both extant species and fossil remains. Some dinosaurs are herbivorous, others carnivorous. Many dinosaurs, including birds, have been bipedal, though many extinct groups were quadrupedal, and some were able to shift between these body postures. Many species possess elaborate display structures such as horns or crests, and some prehistoric groups developed even more elaborate skeletal modifications such as bony armor. Avian dinosaurs have been the planet's dominant flying vertebrate since the extinction of the pterosaurs, and evidence suggests that all ancient dinosaurs built nests and laid eggs much as avian species do today. Although generally known for the large size of some species, most Mesozoic dinosaurs were human-sized or even smaller.
The term "dinosaur" was coined in 1842 by the English paleontologist Richard Owen, and derives from Greek δεινός (deinos) "terrible, powerful, wondrous, potent" + σαῦρος (sauros) "lizard". Through the first half of the 20th century, most of the scientific community believed dinosaurs to have been sluggish, unintelligent cold-blooded animals. Most research conducted since the 1970s, however, has indicated that dinosaurs were active animals with elevated metabolisms and numerous adaptations for social interaction.
Since the first dinosaur fossils were recognized in the early 19th century, mounted fossil dinosaur skeletons or replicas have been major attractions at museums around the world, and dinosaurs have become a part of world culture. They have been featured in best-selling books and films such as Jurassic Park, and new discoveries are regularly covered by the media. In informal speech, the word "dinosaur" is used to describe things that are impractically large, obsolete, or bound for extinction, reflecting the outdated view that dinosaurs were maladapted monsters of the ancient world.

For a long time many scientists thought dinosaurs were polyphyletic with multiple groups of unrelated "dinosaurs" evolving due to similar pressures, but dinosaurs are now known to have formed a single group. Dinosaurs diverged from their archosaur ancestors approximately 230 million years ago during the Middle to Late Triassic period, roughly 20 million years after the Permian–Triassic extinction event wiped out an estimated 95% of all life on EarthRadiometric dating of the rock formation that contained fossils from the early dinosaurgenus Eoraptor establishes its presence in the fossil record at this time. Paleontologists believeEoraptor resembles the common ancestor of all dinosaurs; if this is true, its traits suggest that the first dinosaurs were small, bipedal predators. The discovery of primitive, dinosaur-likeornithodirans such as Marasuchus and Lagerpeton in Argentinian Middle Triassic strata supports this view; analysis of recovered fossils suggests that these animals were indeed small, bipedal predators.
When dinosaurs appeared, terrestrial habitats were occupied by various types of basal archosaurs and therapsids, such as aetosaurs,cynodontsdicynodontsornithosuchidsrauisuchias, and rhynchosaurs. Most of these other animals became extinct in the Triassic, in one of two events. First, at about the boundary between the Carnian and Norian faunal stages (about 215 million years ago), dicynodonts and a variety of basal archosauromorphs, including the prolacertiforms and rhynchosaurs, became extinct. This was followed by the Triassic–Jurassic extinction event (about 200 million years ago), that saw the end of most of the other groups of early archosaurs, like aetosaurs, ornithosuchids, phytosaurs, and rauisuchians. These losses left behind a land fauna of crocodylomorphs, dinosaurs, mammals,pterosaurians, and turtles.
The first few lines of primitive dinosaurs diversified through the Carnian and Norian stages of the Triassic, most likely by occupying the niches of groups that became extinct. Traditionally, dinosaurs were thought to have replaced the variety of other Triassic land animals by proving superior through a long period of competition. This now appears unlikely, for several reasons. Dinosaurs do not show a pattern of steadily increasing in diversity and numbers, as would be predicted if they were competitively replacing other groups; instead, they were very rare through the Carnian, making up only 1–2% of individuals present in faunas. In the Norian, however, after the extinction of several other groups, they became significant components of faunas, representing 50–90% of individuals. Also, what had been viewed as a key adaptation of dinosaurs, their erect stance, is now known to have been present in several contemporaneous groups that were not as successful (aetosaurs, ornithosuchids, rauisuchians, and some groups of crocodylomorphs). Finally, the Late Triassic itself was a time of great upheaval in life, with shifts in plant life, marine life, and climate. Crurotarsans, today represented only by crocodilians but in the Late Triassic also encompassing such now-extinct groups as aetosaurs, phytosaurs, ornithosuchians, and rauisuchians, were actually more diverse in the Late Triassic than dinosaurs, indicating that the survival of dinosaurs had more to do with luck than superiority.
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0 History Of Komodo Dragons


The Komodo dragon (Varanus komodoensis) is a large species of lizard found in theIndonesian islands of KomodoRincaFloresGili Motang and Gili Dasami. A member of the monitor lizard family (Varanidae), it is the largest living species of lizard, growing to a maximum of length 3 metres (9.8 ft) in rare cases and weighing up to around 70 kilograms (150 lb). Their unusual size has been attributed to island gigantism, since there are no other carnivorous animals to fill the niche on the islands where they live.
However, recent research suggests that the large size of komodo dragons may be better understood as representative of a relict population of very large varanid lizards that once lived across Indonesia and Australia, most of which, along with other megafauna, died out after the Pleistocene. Fossils very similar to V. komodoensis have been found in Australia dating to greater than 3.8 million years ago, and its body size remained stable on Flores, one of the handful of Indonesian islands where it is currently found, over the last 900,000 years, "a time marked by major faunal turnovers, extinction of the island's megafauna, and the arrival of early hominids by 880 ka."
As a result of their size, these lizards dominate the ecosystems in which they live. Komodo dragons hunt and ambush prey including invertebratesbirds, and mammals. Their group behaviour in hunting is exceptional in the reptile world. The diet of big Komodo dragons mainly consists of deer, though they also eat considerable amounts of carrion.
Mating begins between May and August, and the eggs are laid in September. About twenty eggs are deposited in abandoned megapode nests or in a self-dug nesting hole. The eggs are incubated for seven to eight months, hatching in April, when insects are most plentiful. Young Komodo dragons are vulnerable and therefore dwell in trees, safe from predators andcannibalistic adults. They take about eight to nine years to mature, and are estimated to live for up to 30 years.
Komodo dragons were first recorded by Western scientists in 1910. Their large size and fearsome reputation make them popular zoo exhibits. In the wild their range has contracted due to human activities and they are listed as vulnerable by the IUC . They are protected under Indonesian law, and a national parkKomodo National Park, was founded to aid protection efforts.

Etymology
The Komodo dragon is also known as the Komodo monitor or the Komodo Island monitor in scientific literature, although this is not very common.[1] To the natives of Komodo Island, it is referred to as orabuaya darat (land crocodile) or biawak raksasa (giant monitor).



Evolutionary history

The evolutionary development of the Komodo dragon started with the Varanus genus, which originated in Asia about 40 million years ago and migrated to Australia. Around 15 million years ago, a collision between Australia and Southeast Asia allowed the varanids to move into what is now the Indonesian archipelago, extending their range as far east as the island of Timor. The Komodo dragon was believed to have differentiated from its Australian ancestors 4 million years ago. However, recent fossil evidence from Queensland suggests that the Komodo dragon evolved in Australia before spreading to Indonesia. Dramatic lowering of sea level during the last glacial period uncovered extensive stretches of continental shelf that the Komodo dragon colonized, becoming isolated in their present island range as sea levels rose afterwards.


Description


In the wild, an adult Komodo dragon usually weighs around 70 kilograms (150 lb), although captive specimens often weigh more. The largest verified wild specimen was 3.13 metres (10.3 ft) long and weighed 166 kilograms (370 lb), including undigested food. The Komodo dragon has a tail as long as its body, as well as about 60 frequently replaced serrated teeth that can measure up to 2.5 cm (1 inch) in length. Its saliva is frequently blood-tinged, because its teeth are almost completely covered by gingival tissue that is naturally lacerated during feeding. This creates an ideal culture for the virulent bacteria that live in its mouth.] It also has a long, yellow, deeply forked tongue.



Reproduction

Mating occurs between May and August, with the eggs laid in September. During this period, males fight over females and territory by grappling with one another upon their hind legs with the loser eventually being pinned to the ground. These males may vomit or defecate when preparing for the fight. The winner of the fight will then flick his long tongue at the female to gain information about her receptivity. Females are antagonistic and resist with their claws and teeth during the early phases of courtship. Therefore, the male must fully restrain the female during coitus to avoid being hurt. Other courtship displays include males rubbing their chins on the female, hard scratches to the back, and licking. Copulation occurs when the male inserts one of his hemipenes into the female's cloaca. Komodo dragons may be monogamous and form "pair bonds", a rare behavior for lizards.

The female lays her eggs in burrows cut into the side of a hill or in the abandoned nesting mounds of the Orange-footed Scrubfowl (a moundbuilder or megapode), with a preference for the abandoned mounds. Clutches contain an average of 20 eggs which have an incubation period of 7–8 months. Hatching is an exhausting effort for the neonates, who break out of their eggshells with an egg tooth that falls off soon after. After cutting out the hatchlings may lie in their eggshells for hours before starting to dig out of the nest. They are born quite defenseless, and many are eaten by predators.
Young Komodo dragons spend much of their first few years in trees, where they are relatively safe from predators, including cannibalistic adults, who make juvenile dragons 10% of their diet. According to David Attenborough, the habit of cannibalism may be advantageous in sustaining the large size of adults, as medium-sized prey on the islands is rare. When the young must approach a kill, they roll around in fecal matter and rest in the intestines of eviscerated animals to deter these hungry adults. Komodo dragons take about three to five years to mature, and may live for up to 50 years.


Parthenogenesis


A Komodo dragon at London Zoo named Sungai laid a clutch of eggs in late 2005 after being separated from male company for more than two years. Scientists initially assumed that she had been able to store sperm from her earlier encounter with a male, an adaptation known assuperfecundation. On December 20, 2006, it was reported that Flora, a captive Komodo dragon living in the Chester Zoo in England, was the second known Komodo dragon to have laid unfertilized eggs: she laid 11 eggs, and 7 of them hatched, all of them male. Scientists atLiverpool University in England performed genetic tests on three eggs that collapsed after being moved to an incubator, and verified that Flora had never been in physical contact with a male dragon. After Flora's eggs' condition had been discovered, testing showed that Sungai's eggs were also produced without outside fertilization. On January 31, 2008, the Sedgwick County Zoo inWichita, Kansas became the first zoo in the Americas to document parthenogenesis in Komodo dragons. The zoo has two adult female Komodo dragons, one of which laid about 17 eggs on May 19–20, 2007. Only two eggs were incubated and hatched due to space issues; the first hatched on January 31, 2008 while the second hatched on February 1. Both hatchlings were males.
Komodo dragons have the ZW chromosomal sex-determination system, as opposed to the mammalian XY system. Male progeny prove that Flora's unfertilized eggs were haploid (n) and doubled their chromosomes later to become diploid (2n) (by being fertilized by a polar body, or by chromosome duplication without cell division), rather than by her laying diploid eggs by one of the meiosis reduction-divisions in herovaries failing. When a female Komodo dragon (with ZW sex chromosomes) reproduces in this manner, she provides her progeny with only one chromosome from each of her pairs of chromosomes, including only one of her two sex chromosomes. This single set of chromosomes is duplicated in the egg, which develops parthenogenetically. Eggs receiving a Z chromosome become ZZ (male); those receiving a W chromosome become WW and fail to develop.

It has been hypothesized that this reproductive adaptation allows a single female to enter an isolated ecological niche (such as an island) and by parthenogenesis produce male offspring, thereby establishing a sexually reproducing population (via reproduction with her offspring that can result in both male and female young). Despite the advantages of such an adaptation, zoos are cautioned that parthenogenesis may be detrimental to genetic diversity.
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